What's the difference between pule and puler?

Pule

Definition:

(v. i.) To cry like a chicken.
(v. i.) To whimper; to whine, as a complaining child.

Example Sentences:

(1) The previously uncharacterized third and fourth genes (pulE and pulF) of the pullulanase secretion gene operon of Klebsiella oxytoca strain UNF5023 are, respectively, predicted to encode a 55 kDa polypeptide with a putative nucleotide-binding site, and a highly hydrophobic 44 kDa polypeptide that probably spans the cytoplasmic membrane several times.
(2) Under basal conditions oral penbutolol induced a decrease of pule rate and blood pressure but no change in plasma or urinary catecholamines.
(3) This suppression was due to increased [K+]0 and not to K-induced depolarization because it persisted when membrane potential was held by means of a conditioning hyperpolarizing puled gradually after maximum repolarization.
(4) 25 laryngeal carcinomas were investigated by pules cytophotometry, stained with ethidium bromide after pepsination.
(5) These results are in line with the predicted absence from PulE of a region of sufficient hydrophobicity to function as a signal sequence.
(6) Expression of pulE in minicells or under the control of a strong bacteriophage T7 promoter resulted in the production of a c. 58 kDa cytoplasmic protein.
(7) Gene disruption experiments indicated that both pulE and pulF are required for pullulanase secretion in Escherichia coli K-12.
(8) A representative PulE-beta-galactosidase hybrid protein created by Tnlac mutagenesis was also found mainly in the cytoplasm.
(9) These include the P. aeruginosa PilB protein, the ComG ORF-1 protein from the Bacillus subtilis comG operon (necessary for competence), the PulE protein from the Klebsiella oxytoca (formerly K. pneumoniae) pulC-O operon (involved in pullulanase export), and the VirB-11 protein from the virB operon (involved in virulence) which is located on the Agrobacterium tumefaciens Ti plasmid.
(10) They issue great puling statements about income imbalance in a game that pays them $100m per annum just for the act of being able to cash checks and maybe pay attention long enough to run a franchise into the ground.
(11) In addition, PulE protein has consensus sequences found in a wide variety of nucleotide-binding proteins.
(12) The deduced amino acid sequence of ORF1 is related to the Klebsiella pneumoniae PulE protein, to the Bacillus subtilis ComG ORF1 and to the Agrobacterium tumefaciens VirB ORF11 products.